Lesser Black-backed Gull (graellsii & intermedius)

(last update: 09-06-2013)

Home

LBBG rings

lbbg 1cy May
lbbg 1cy June
lbbg 1cy July
lbbg 1cy August
lbbg 1cy September
lbbg 1cy October
lbbg 1cy Nov - Dec

lbbg 2cy Jan-Febr
lbbg 2cy Mar-April
lbbg 2cy May
lbbg 2cy June
lbbg 2cy July
lbbg 2cy August
lbbg 2cy September
lbbg 2cy October
lbbg 2cy Nov - Dec

lbbg 3cy Jan-April
lbbg 3cy May
lbbg 3cy June
lbbg 3cy July
lbbg 3cy August
lbbg 3cy September
lbbg 3cy October
lbbg 3cy Nov - Dec

lbbg sub-ad Jan-April
lbbg sub-ad May
lbbg sub-ad June
lbbg sub-ad July
lbbg sub-ad Aug
lbbg sub-ad Sept
lbbg sub-ad Oct
lbbg sub-ad Nov - Dec

lbbg ad January
lbbg adult February
lbbg adult Mar-April
lbbg adult May
lbbg adult June
lbbg adult July
lbbg ad August
lbbg ad September
lbbg ad October
lbbg ad Nov - Dec


third calendar year: June

According to most field guides, second summer (third calendar year) LBBG normally show a grey saddle (adult-like feathers on mantle and scapulars), brown wing-coverts, 2nd generation primaries and secondaries, and a 2nd generation tail showing a broad tail-band.

However, in the field, almost all 3CY June birds have some of the coverts moulted into third generation. Sometimes the entire median covert row has been replaced, creating a grey bar.

But quite some birds have replaced more than a median covert row; they also replaced tail-feahers, or sometimes even secondaries and primaries. Especially these advanced birds appear much more adult-like than text-book 3CY birds.

Overall, there seems to be some variation in birds of this age class. The feathers showing most variation are the rectrices; it might be anything from a clear tail-band, spaghetti pattern, black-and-white variegated pattern to a complete snow white tail (although close examination often reveals small black spots near the feather centre). When secondaries have been renewed in winter, they normally stand out as adult-like grey feathers with white tips. Most of these advanced individuals seem to be slightly darker and more elongated; they probably originate from intermedius populations. Note however that a bird ringed in Norway has been recovered in the Maasvlakte colony, where it actually bred. Dutch colour rings have been found in Scandinavia in reverse.

Ageing of 3CY birds in spring

3CY birds in spring (at their end of 2nd winter plumage) can be highly variable. Much has to do with the extent of previous moult cycles, and in particular the timing of these moults. Classic Lesser Black-backed Gulls "graellsii-type birds" very much resemble Herring Gull in moult timing and extent. Scandinavian intermedius-type birds often show a shifted moult timing and replace feathers in late-winter. As a consequence, the patterns of these replaced feathers may be different (note that this is largely due to moult TIMING, and in fact we look at same generatrion feathers).

graellsii-type LBBG

3CY Herring Gull and 3CY graellsii-type LBBG in spring look very similar in general. The scapular and mantle are adult-like grey wing-coverts are barred/brown, the 2nd generation tail show a broad band, 2nd generation secondaries have dark brownish centres and inner primaries are not adult-like but still have a brownish wash and only very ill-defined diffuse dark tips. Second generation P4-P5 have no sharply defined sub-terminal band plus white tip (as this is a good pointer to 3rd generation feathers and hence such birds are in 4CY).

Above: 3CY Herring Gull, Katwijk, the Netherlands, May 19 2013. Typical 3cy bird with 2nd generation primaries, secondaries and tail-feathers. Outer primaries brown without sharply defined white tips. Inner primaries P3-P5 without black sub-terminal band, these 2nd generation inner primaries only show a diffuse dark tip and outerweb. Secondaries with brownish barred and vermiculated centres. All tail-feathers with dark distal part.

Below: Classic 3CY graellsii in June with adult-like grey mantle + scapulars (grey saddle) well demarcated from brown coverts. P5-P10 old 2nd generation feathers, P5 with diffuse brown outerweb, but no obvious blackish sub-terminal band. Second generation tail.

Ageing 3CY birds from 2CY birds is often rather straightforward. However, 3CY and 4CY birds may look similar. Typical 3CY graellsii-type LBBG have 2nd generation P4-P5, which has a diffuse brown tip and outerweb. 4CY are one year older, and the inner primaries are reminiscent of adults: P4-P5 normally have sharply defined blackish sub-terminal patterns and the innerwebs are greysih, not brownish.

Above: 4CY graellsii in June with adult-like upperparts and coverts. P3 missing, P4-P10 old 3rd generation feathers, typically P4-P5 with obvious blackish sub-terminal bands and white tips. Third generation tail and secondaries (note bold white tips on secondaries, combined with blackish immature centres).

intermedius-type LBBG

3CY intermedius-type LBBG in spring may look very similar to older birds, and sometimes is really difficult to tell them apart. Best clue, again, are the central primaries P3-P5, in combination with patterns on the secondaries.
The main problem with intermedius birds is, they often have a late-winter moult which can be very extensive, including all tail-feathers and many secondaries. Some birds even replace primaries on the winter grounds, prior to flying north again. Such birds adopt a fuscus-like moult strategy, with old outer and new inner primaries in spring and its only possible to ID these intermedius birds by the paler upperparts or/and ring, if present.

Above: 3CY intermedius in June with adult-like upperparts and coverts. P2 missing, P3-P10 old 2nd generation feathers, typically with diffuse brown outerweb, and no obvious blackish sub-terminal band. Advanced moult on winter grounds couple of months ago: inner secondaries and all tail-feathers replaced, still looking fresh.
Below: 3CY intermedius in June with adult-like upperparts and coverts. P3 missing, P4 replaced 3rd gen feather, P5-P10 old 2nd generation feathers, typically with diffuse brown outerweb, and no obvious blackish sub-terminal band. Advanced moult on winter grounds couple of months ago: 'jumping' moult sequence in secondaries and all tail-feathers replaced, new feathers still looking fresh.

Detailed description of  3cy LBBG in June

Mantle and scapulars

By June, adult-like grey feathers replace brown-hued old scapulars, probably at least for the third time, resulting in a fourth generation scapular region. Some old scapulars show a faint anchor pattern on a brown base.

Above: graellsii L-97134, ringed as pullus at Zeebrugge, Belgium on July 09 2001. Very immature in upper-parts pattern, unlike the adult-grey scapulars in most 3cy LBBG.
Below: Text-book 3CY graellsii in June with adult-like grey mantle + scapulars (grey saddle) well demarcated from brown coverts.

Moult of coverts

The section dealing with 2CY autumn birds already show that a subsequent moult may follow in quite some birds in autumn, called a "partial autumn moult". Normally this will involve upper tertials (TT), inner greater coverts (GC), many of the inner median and lower lesser coverts (MC and LLC).

Above: 2CY September graellsii blue DUAK, ringed as pullus in Belgium, July 07 2006. Partial autumn moult obvious in MC and upper TT. Also GC1 missing and few LLC growing.
Below: 3CY January graellsii blue HHAS, ringed as pullus in Belgium in 2008. Those wing-coverts replaced in the partial autumn moult remain obvious throughout winter period.
Above: 3CY June LBBG. Central TT and most rectrices replaced recently, probably late winter March-April. By June, these feathers stand out as very adult-like, with neat fringes and no bleached brownish hue. Without winter observations of a particular individual, it remains speculation if adult-like coverts were replaced last autumn or in late winter. Complete moult not started: P1 still present.
Below: even more advanced late winter moult also included secondaries. Very fresh looking coverts probably also replaced recently, but late winter observations of this individual necessary to clarify this.

Late winter (March-April) occasionally there seems to be a moult taken place in some birds, especially intermedius. This moult has not the common sequence we witness in summer, the complete moult. In summer, complete moult starts with dropping P1 and P2, then followed by other primaries and simultaneously outer median coverts (outer MC) are dropped. Another moult wave in wing-coverts starts at the very innermost feathers, and two waves evolve through the wing-covert panel.
However, few 3CY intermedius in late winter, March-April, may start replacing wing-coverts, rectrices and even secondaries or primaries. When these birds turn up in NW Europe in June, these late-winter feathers still appear very fresh and neat.

Above: 3CY June LBBG. Complete moult commenced, inner primaries dropped, and simultaneously the outer MC have been shed as well (arrow A). It is normal to skip fresh feathers (from partial autumn moult) in the early moult stage, hence first replace very old feathers.

By June, 3CY LBBG start moulting the median coverts, starting with outermost feathers when P1-P3 are dropped. New third generation coverts are easily recognized as grey adult-like feathers.

Tail-feather (rectrices) moult

The exact rectrices-moult is often difficult to ascertain. In graellsii, most common pattern at this age is a worn second generation tail, i.e. a tail with black marks or tail feathers with a broad black tail-band. This is the classic 2nd generation tail pattern for spring 3CY birds.

Above: 3CY June LBBG. Classic tail, old, slightly worn 2nd generation rectrices.
Below: 3CY June LBBG. Some rectrices old 2nd generation, but some replaced, now adult-like white.
Below: 3CY June LBBG. All rectrices replaced, now adult-like white.

June is early in the complete moult cycle, so only very few birds are actively moulting tail-feathers, starting with the innermost pair (R1, these are the feathers most exposed to sun and weather conditions). More than 30% of 3cy LBBGs has moulted one or more pairs of tail-feathers last winter, resulting in a blocked pattern of black and white feathers or even a completely white tail. This winter moult in the tail involved rectrices randomly, creating an unpredictable black and white tail in spring.

Remiges moult

At the start of June some birds have already shed P1 in the complete moult, while at the end of June most birds have P2 replaced and P4 shed. New third generation primaries are very adult-like (adult-grey with white tip) and differ clearly from the plain black-brown second generation primaries. 
The differences with real adult primaries can be found in the tips which are normally smaller in third generation primaries, P9 normally lacks a mirror, P10 has a smaller mirror and there is an extensive amount of black on P3 and P4 (more than on pure adult primaries and it's not unusual to find a complete sub-terminal band on P4 in third generation primaries, where most adults have the black markings on P4 confined to the outer web). 
Often the third generation outer primaries show a brown hue, another sign of immaturity (but these features don't apply on June LBBG of course: P9 and P10 are full-grown in November). Older birds show more blackish P5-P10 (the age effects the feathers: old feathers are stronger, making the effect of bleaching and wear less obvious).

By June, the old second generation P10 is half a year of age and shows an obvious white shaft, creating a broad white line in a pale brown feather, on the underside of p10. There is no active moult in the secondaries yet.

As already mentioned under "Moult of Coverts", some 3CY LBBG return from the winter grounds with replaced (3rd generation) secondaries or even primaries. Example birds on this page illustrate this, and we believe this to occur more often in intermedius than it does in graellsii.

Above: 3CY June LBBG. Complete moult not started, P1 still present. However, complete set of rectrices plus secondaries and several primaries clearly 3rd generation.

Tables:

Primary moult score in 3cy LBBG at Maasvlakte, the Netherlands, June 17 2002. Total n: 25
p1 old 1
p2 old 4
p3 old 6
p4 old 9
p5 old 5

LBBG 3cy Belgium: graellsii


flying LBBG:

3cy graellsii like birds.

3cy graellsii LBBG in June, ringed in the Netherlands. (135385 bytes)LBBG TWAW 3cy June 16 2013, Noordwijk, the Netherlands. Picture: Frank van der Meer & Ruud Altenburg. LBBG 3cy June 09 2013, Noordwijk, the Netherlands. Classic 3cy LBBG with 2nd generation flight feathers. 


LBBG 3cy the Netherlands: graellsii / Dutch intergrade

 

3cy graellsii LBBG in June, ringed in the Netherlands. (135385 bytes)LBBG 3cy June 24 2013, Noordwijk, the Netherlands.
sub-adult LBBG in August, ringed in Belgium. (82012 bytes)LBBG 3-4cy YK June 2009 & August 2010, Barneveld, The Netherlands. LBBG 3cy June 10 2013, Noordwijk, the Netherlands.
3cy graellsii LBBG in June, ringed in the Netherlands. (92536 bytes)LBBG 3cy S9 June 24 2009, Barneveld, the Netherlands. LBBG 3cy June 10 2013, Noordwijk, the Netherlands.
sub-adult LBBG in August, ringed in Belgium. (82012 bytes)LBBG 3cy D0 June 10 2010, Utrecht, The Netherlands. Picture: Herman Bouman. LBBG 3cy June 10 2013, Noordwijk, the Netherlands.
3cy graellsii LBBG in June, ringed in the Netherlands. (135385 bytes)LBBG 0* 3cy June 24 2013, Noordwijk, the
Netherlands. Classic 3cy LBBG, complete moult started.
3cy graellsii LBBG in June, ringed in the Netherlands. (135385 bytes)LBBG 3cy June 24 2013, Noordwijk, the Netherlands.
3cy graellsii LBBG in June, ringed in the Netherlands. (135385 bytes)LBBG 6N 3cy June 16 2013, Noordwijk, the Netherlands. Picture: Ruud Altenburg. LBBG 3cy June 10 2013, Noordwijk, the Netherlands.
3cy graellsii LBBG in June, ringed in the Netherlands. (135385 bytes)LBBG Y2 3cy June 16 2013, Noordwijk, the
Netherlands.Picture: Frank van der Meer.
LBBG 3cy June 10 2013, Noordwijk, the Netherlands.
3cy graellsii LBBG in June, ringed in the Netherlands. (135385 bytes)LBBG 13 3cy June 16 2013, Noordwijk, the Netherlands. Picture: Frank van der Meer. LBBG 3cy June 10 2013, Noordwijk, the Netherlands.
3cy graellsii LBBG in June, ringed in the Netherlands. (135385 bytes)LBBG 1C 3cy June 16 2013, Noordwijk, the Netherlands. Picture: Ruud Altenburg. LBBG 3cy June 09 2013, Noordwijk, the Netherlands. Classic 3cy LBBG, complete moult started.
3cy graellsii LBBG in June, ringed in the Netherlands. (135385 bytes)LBBG 1Y 3cy June 16 2013, Noordwijk, the Netherlands. Picture: Ruud Altenburg. LBBG 3cy June 10 2013, Noordwijk, the Netherlands.
3cy graellsii LBBG in June, ringed in the Netherlands. (135385 bytes)LBBG 27 3cy June 16 2013, Noordwijk, the Netherlands. Picture: Frank van der Meer. LBBG 3cy June 10 2013, Noordwijk, the Netherlands.
3cy graellsii LBBG in June, ringed in the Netherlands. (135385 bytes)LBBG 49 3cy June 09 2013, Noordwijk, the Netherlands. Classic 3cy LBBG, complete moult started. LBBG 3cy June 10 2013, Noordwijk, the Netherlands.
3cy graellsii LBBG in June, ringed in the Netherlands. (135385 bytes)LBBG KAKP 3cy June 24 2013, Noordwijk, the
Netherlands. Classic 3cy LBBG, complete moult started.

LBBG 3cy June 10 2013, Noordwijk, the Netherlands.

3cy graellsii LBBG in June, ringed in the Netherlands. (135385 bytes)LBBG KALF 3cy June 16 2013, Noordwijk, the Netherlands. Picture: Ruud Altenburg.

LBBG 3cy June 10 2013, Noordwijk, the Netherlands.

3cy graellsii LBBG in June, ringed in the Netherlands. (135385 bytes)LBBG KAMU 3cy June 16 2013, Noordwijk, the Netherlands. Picture: Ruud Altenburg. LBBG 3cy June 10 2013, Noordwijk, the Netherlands.
3cy graellsii LBBG in June, ringed in the Netherlands. (135385 bytes)LBBG YAWA 3cy June 16 2013, Noordwijk, the Netherlands. Picture: Ruud Altenburg. 3cy graellsii LBBG in June, ringed in the Netherlands. (135385 bytes)LBBG 3cy June 24 2013, Noordwijk, the Netherlands.
3cy graellsii LBBG in June, ringed in the Netherlands. (135385 bytes)LBBG YAWZ 3cy June 16-22 2013, Noordwijk, the Netherlands. Picture: Ruud Altenburg & Maarten van Kleinwee. Tail and several secondaries replaced at winter grounds. LBBG 3cy June 10 2013, Noordwijk, the Netherlands.
3cy graellsii LBBG in June, ringed in the Netherlands. (135385 bytes)LBBG YAXS 3cy June 24 2013, Noordwijk, the
Netherlands. Classic 3cy LBBG, complete moult started.
LBBG 3cy June 09 2013, Noordwijk, the Netherlands. Classic 3cy LBBG, complete moult started.
3cy graellsii LBBG in June, ringed in the Netherlands. (135385 bytes)LBBG YAZP 3cy June 09 2013, Noordwijk, the
Netherlands. Classic 3cy LBBG, complete moult started.
LBBG 3cy June 10 2013, Noordwijk, the Netherlands.
3cy graellsii LBBG in June, ringed in the Netherlands. (100728 bytes)LBBG C00 3cy June 18 2005, Amsterdam, the Netherlands. Picture Ruud Altenburg. LBBG 3cy June 10 2013, Noordwijk, the Netherlands.
3cy graellsii LBBG in June, ringed in the Netherlands. (92536 bytes)LBBG C73 3cy June 18 2005, Amsterdam, the Netherlands. Picture Ruud Altenburg. LBBG 3cy June 10 2013, Noordwijk, the Netherlands.
3cy graellsii LBBG in June, ringed in the Netherlands. (135385 bytes)LBBG 5.412.031 3cy June 12 2005, Amsterdam, the Netherlands. Picture Ruud Altenburg. LBBG 3cy June 10 2013, Noordwijk, the Netherlands.
3cy graellsii LBBG in June, ringed in the Netherlands. (135385 bytes)LBBG 5.412.031 3cy June 12 2005, Amsterdam, the Netherlands. Picture Ruud Altenburg. LBBG 3cy June 10 2013, Noordwijk, the Netherlands.
3cy graellsii LBBG in June, ringed in the Netherlands. (88149 bytes)LBBG 5.412.128 3cy June 25 2005, Amsterdam, the Netherlands. Picture Ruud Altenburg. LBBG 3cy June 10 2013, Noordwijk, the Netherlands.

3cy graellsii LBBG in June, ringed in the Netherlands. (85566 bytes)LBBG 5.412.152 3cy June 12 2005, Amsterdam, the Netherlands. Picture Ruud Altenburg.

LBBG 3cy June 10 2013, Noordwijk, the Netherlands.
3cy graellsii LBBG in June, ringed in the Netherlands. (85566 bytes)LBBG 5.414.319 3cy June 05 2006, Amsterdam, the Netherlands. Picture Ruud Altenburg. LBBG 3cy June 10 2013, Noordwijk, the Netherlands.
3cy graellsii LBBG in June, ringed in the Netherlands. (85566 bytes)LBBG E 3cy June 06 2006, Amsterdam, the Netherlands. Picture Ruud Altenburg. LBBG 3cy June 10 2013, Noordwijk, the Netherlands.


LBBG 3cy Germany: graellsii-intermedius transition

LBBG 3cy June 09 2013, Noordwijk, the Netherlands. Classic 3cy LBBG, complete moult started.

sub-adult LBBG in August, ringed in Belgium. (82012 bytes)LBBG 3cy H018 June 23 2009, Utrecht, The Netherlands. Picture: Herman Bouman.

LBBG 3cy June 10 2013, Noordwijk, the Netherlands.
adult LBBG Dutch intergrade in September, ringed in the Netherlands. (85708 bytes)LBBG Dutch intergrade H501 Seen as 3cy and 4cy, Leiden, the Netherlands. Pictures: Maarten van Kleinwee. LBBG 3cy June 10 2013, Noordwijk, the Netherlands.
3cy graellsii LBBG in June, ringed in the Netherlands. (135385 bytes)LBBG 3cy HN46 June 09 2013, Noordwijk, the Netherlands. Classic 3cy LBBG, complete moult
started.
3cy graellsii LBBG in June, ringed in the Netherlands. (135385 bytes)LBBG 3cy June 24 2013, Noordwijk, the Netherlands.

LBBG 3cy Norway: intermedius
3cy graellsii LBBG in June, ringed in the Netherlands. (135385 bytes)LBBG 3cy June 24 2013, Noordwijk, the Netherlands.
3cy graellsii LBBG in June, ringed in the Netherlands. (135385 bytes)LBBG JJ1A 3cy June 24 2013, Noordwijk, the
Netherlands. Classic 3cy LBBG, complete moult started.
3cy graellsii LBBG in June, ringed in the Netherlands. (135385 bytes)LBBG 3cy June 24 2013, Noordwijk, the Netherlands.
3cy graellsii LBBG in June, ringed in the Netherlands. (135385 bytes)LBBG JJ5H 3cy June 16 2013, Noordwijk, the
Netherlands. Picture: Ruud Altenburg.
3cy graellsii LBBG in June, ringed in the Netherlands. (135385 bytes)LBBG 3cy June 24 2013, Noordwijk, the Netherlands.
3cy intermedius in June, ringed in Norway. (64435 bytes)LBBG J1E7 3cy, June 24 2003, IJmuiden, the Netherlands. Ringed in Norway. 3cy graellsii LBBG in June, ringed in the Netherlands. (135385 bytes)LBBG 3cy June 24 2013, Noordwijk, the Netherlands.


LBBG 3cy Spain: unknown - ringed in winter

 

3cy graellsii LBBG in June, ringed in the Netherlands. (135385 bytes)LBBG 3cy June 24 2013, Noordwijk, the Netherlands.
3cy graellsii LBBG in June, ringed in the Netherlands. (135385 bytes)LBBG N04U 3cy June 16 2013, Noordwijk, the Netherlands.Picture: Frank van der Meer & Ruud Altenburg. 3cy graellsii LBBG in June, ringed in the Netherlands. (135385 bytes)LBBG 3cy June 24 2013, Noordwijk, the Netherlands.

3CY LBBG

with arrested primary moult

3cy graellsii LBBG in June, ringed in the Netherlands. (135385 bytes)LBBG 3cy June 24 2013, Noordwijk, the Netherlands.
2261.jpg (50391 bytes)LBBG intermedius 3cy June 28 2003, Dannes - Boulogne/Mer, NW FRance. P1 4th generation, P2-P3 missing, P4-P7 3rd generation, P8-P10 old 2nd generation.  3cy graellsii LBBG in June, ringed in the Netherlands. (135385 bytes)LBBG 3cy June 24 2013, Noordwijk, the Netherlands.
LBBG intermedius(?) 3cy June 30 2001, IJmuiden, the Netherlands. Very advanced 2nd summer: almost all upper parts moulted and arrested moult in the primaries.  3cy graellsii LBBG in June, ringed in the Netherlands. (135385 bytes)LBBG 3cy June 24 2013, Noordwijk, the Netherlands.
LBBG 3cy June 10 2013, Noordwijk, the Netherlands. Fuscus-like bird. 3cy graellsii LBBG in June, ringed in the Netherlands. (135385 bytes)LBBG 3cy June 24 2013, Noordwijk, the Netherlands.
LBBG 3cy June 23 2006, Amsterdeam, the Netherlands. Arrested moult in primaries.  3cy graellsii LBBG in June, ringed in the Netherlands. (135385 bytes)LBBG 3cy June 24 2013, Noordwijk, the Netherlands.
LBBG 3cy June 09 2013, Noordwijk, the Netherlands. Advanced 3cy LBBG, arrested moult in primaries.  3cy graellsii LBBG in June, ringed in the Netherlands. (135385 bytes)LBBG 3cy June 24 2013, Noordwijk, the Netherlands.
LBBG 3cy June 09 2013, Noordwijk, the Netherlands. Advanced 3cy LBBG, arrested moult in primaries.  LBBG 3cy June 10 2013, Noordwijk, the Netherlands.
LBBG 3cy June 09 2013, Noordwijk, the Netherlands. Advanced 3cy LBBG, arrested moult in primaries. 

flying LBBG:

3cy birds with replaced tail-feathers.

LBBG 3cy June 09 2013, Noordwijk, the Netherlands. Advanced 3cy LBBG, arrested moult in primaries.  LBBG 3cy June 10 2013, Noordwijk, the Netherlands.
LBBG 3cy June 09 2013, Noordwijk, the Netherlands. Advanced 3cy LBBG, arrested moult in primaries.  3cy graellsii LBBG in June, ringed in the Netherlands. (135385 bytes)LBBG 3cy June 24 2013, Noordwijk, the Netherlands.
3cy graellsii LBBG in June, ringed in the Netherlands. (135385 bytes)LBBG 3cy June 24 2013, Noordwijk, the Netherlands. 3cy graellsii LBBG in June, ringed in the Netherlands. (135385 bytes)LBBG 3cy June 24 2013, Noordwijk, the Netherlands.
3cy graellsii LBBG in June, ringed in the Netherlands. (135385 bytes)LBBG 3cy June 24 2013, Noordwijk, the Netherlands. 3cy graellsii LBBG in June, ringed in the Netherlands. (135385 bytes)LBBG 3cy June 24 2013, Noordwijk, the Netherlands.
LBBG 3cy June 10 2013, Noordwijk, the Netherlands. Advanced 3cy LBBG, arrested moult in primaries.  LBBG 3cy June 10 2013, Noordwijk, the Netherlands.
LBBG 3cy June 10 2013, Noordwijk, the Netherlands. Left wing: P5-P6 replaced. Right wing: P1?, P3-P5 replaced. LBBG 3cy June 10 2013, Noordwijk, the Netherlands.
LBBG 3cy June 10 2013, Noordwijk, the Netherlands. P1 replaced. LBBG 3cy June 10 2013, Noordwijk, the Netherlands.
3cy graellsii LBBG in June, ringed in the Netherlands. (135385 bytes)LBBG 3cy June 24 2013, Noordwijk, the
Netherlands. P5 replaced on winter grounds; complete moult started.
LBBG 3cy June 10 2013, Noordwijk, the Netherlands.
3cy graellsii LBBG in June, ringed in the Netherlands. (135385 bytes)LBBG 3cy June 24 2013, Noordwijk, the
Netherlands. P4-P6 replaced on winter grounds; complete moult started.
LBBG 3cy June 10 2013, Noordwijk, the Netherlands.
3cy graellsii LBBG in June, ringed in the Netherlands. (135385 bytes)LBBG 3cy June 24 2013, Noordwijk, the Netherlands. P4 replaced on winter grounds; complete moult started. LBBG 3cy June 10 2013, Noordwijk, the Netherlands.

flying LBBG:

3cy birds with replaced secondaries.

LBBG 3cy June 10 2013, Noordwijk, the Netherlands.
3cy graellsii LBBG in June, ringed in the Netherlands. (135385 bytes)LBBG 3cy June 24 2013, Noordwijk, the Netherlands. LBBG 3cy June 10 2013, Noordwijk, the Netherlands.
LBBG 3cy June 10 2013, Noordwijk, the Netherlands. 3cy graellsii LBBG in June, ringed in the Netherlands. (135385 bytes)LBBG 3cy June 24 2013, Noordwijk, the Netherlands.
LBBG 3cy June 10 2013, Noordwijk, the Netherlands. 3cy graellsii LBBG in June, ringed in the Netherlands. (135385 bytes)LBBG 3cy June 24 2013, Noordwijk, the Netherlands.
LBBG 3cy June 10 2013, Noordwijk, the Netherlands. LBBG 3cy June 10 2013, Noordwijk, the Netherlands.
LBBG 3cy June 10 2013, Noordwijk, the Netherlands. 3cy graellsii LBBG in June, ringed in the Netherlands. (135385 bytes)LBBG 3cy June 24 2013, Noordwijk, the Netherlands.
LBBG 3cy June 10 2013, Noordwijk, the Netherlands. 3cy graellsii LBBG in June, ringed in the Netherlands. (135385 bytes)LBBG 3cy June 24 2013, Noordwijk, the Netherlands.
LBBG 3cy June 10 2013, Noordwijk, the Netherlands. 3cy graellsii LBBG in June, ringed in the Netherlands. (135385 bytes)LBBG 3cy June 24 2013, Noordwijk, the Netherlands.
LBBG 3cy June 10 2013, Noordwijk, the Netherlands. LBBG 3cy June 10 2013, Noordwijk, the Netherlands.
LBBG 3cy June 10 2013, Noordwijk, the Netherlands. LBBG 3cy June 10 2013, Noordwijk, the Netherlands.
3cy graellsii LBBG in June, ringed in the Netherlands. (135385 bytes)LBBG 3cy June 24 2013, Noordwijk, the Netherlands. LBBG 3cy June 10 2013, Noordwijk, the Netherlands.
  3cy graellsii LBBG in June, ringed in the Netherlands. (135385 bytes)LBBG 3cy June 24 2013, Noordwijk, the Netherlands.
  3cy graellsii LBBG in June, ringed in the Netherlands. (135385 bytes)LBBG 3cy June 24 2013, Noordwijk, the Netherlands.
  LBBG 3cy June 10 2013, Noordwijk, the Netherlands.
  LBBG 3cy June 10 2013, Noordwijk, the Netherlands.
  LBBG 3cy June 10 2013, Noordwijk, the Netherlands.
  LBBG sub-adult / 3cy June 10 2013, Noordwijk, the Netherlands.
  LBBG 3cy June 10 2013, Noordwijk, the Netherlands.
  LBBG 3cy June 10 2013, Noordwijk, the Netherlands.
  LBBG 3cy June 10 2013, Noordwijk, the Netherlands.
  LBBG 3cy June 10 2013, Noordwijk, the Netherlands.
  LBBG 3cy June 10 2013, Noordwijk, the Netherlands.
  LBBG 3cy June 10 2013, Noordwijk, the Netherlands.

 

LBBG 3cy June 10 2013, Noordwijk, the Netherlands.
  LBBG 3cy June 10 2013, Noordwijk, the Netherlands.
  LBBG 3cy June 10 2013, Noordwijk, the Netherlands.
  LBBG 3cy June 10 2013, Noordwijk, the Netherlands.
  LBBG 3cy June 10 2013, Noordwijk, the Netherlands.
  LBBG 3cy June 10 2013, Noordwijk, the Netherlands.
  LBBG 3cy June 10 2013, Noordwijk, the Netherlands.
  3cy graellsii LBBG in June, ringed in the Netherlands. (135385 bytes)LBBG 3cy June 24 2013, Noordwijk, the Netherlands.
  3cy graellsii LBBG in June, ringed in the Netherlands. (135385 bytes)LBBG 3cy June 24 2013, Noordwijk, the Netherlands.
  3cy graellsii LBBG in June, ringed in the Netherlands. (135385 bytes)LBBG 3cy June 24 2013, Noordwijk, the Netherlands.
  3cy graellsii LBBG in June, ringed in the Netherlands. (135385 bytes)LBBG 3cy June 24 2013, Noordwijk, the Netherlands.
  3cy graellsii LBBG in June, ringed in the Netherlands. (135385 bytes)LBBG 3cy June 24 2013, Noordwijk, the Netherlands.
  3cy graellsii LBBG in June, ringed in the Netherlands. (135385 bytes)LBBG 3cy June 24 2013, Noordwijk, the Netherlands.
  LBBG 3cy June 09 2013, Noordwijk, the Netherlands. Classic 3cy LBBG, complete moult not started.