Lesser Black-backed Gull- Kleine Mantelmeeuw (graellsii & intermedius)

(last update: 30 April 2014)

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LBBG rings

lbbg 1cy May
lbbg 1cy June
lbbg 1cy July
lbbg 1cy August
lbbg 1cy September
lbbg 1cy October
lbbg 1cy Nov - Dec

lbbg 2cy Jan-Febr
lbbg 2cy Mar-April
lbbg 2cy May
lbbg 2cy June
lbbg 2cy July
lbbg 2cy August
lbbg 2cy September
lbbg 2cy October
lbbg 2cy Nov - Dec

lbbg 3cy Jan-April
lbbg 3cy May
lbbg 3cy June
lbbg 3cy July
lbbg 3cy August
lbbg 3cy September
lbbg 3cy October
lbbg 3cy Nov - Dec

lbbg sub-ad Jan-April
lbbg sub-ad May
lbbg sub-ad June
lbbg sub-ad July
lbbg sub-ad Aug
lbbg sub-ad Sept
lbbg sub-ad Oct
lbbg sub-ad Nov - Dec

lbbg ad January
lbbg adult February
lbbg adult Mar-April
lbbg adult May
lbbg adult June
lbbg adult July
lbbg ad August
lbbg ad September
lbbg ad October
lbbg ad Nov - Dec

adult: October

Movements

By October, all LBBG have left the colonies and surrounding feeding grounds near the Maasvlakte (the Netherlands) and migrate south towards France, Portugal and Morocco. In November and to a lesser extend December, a couple of hundreds remain on continental NW Europe, as north as the south of the Netherlands along the coast at Westkapelle, 65 km south of the Maasvlakte (see map). As it seems, every year a larger group of LBBGs stay "at the most northern latitude as possible", i.e. migrate south only when winter conditions force them to.

By late November 2000 (a rather mild winter) ringed individuals from Scandinavia, Denmark, the Netherlands, Belgium and the U.K. were present at Westkapelle. When winter really started in December 2000, this group moved south as well to unknown wintering grounds, probably the Atlantic coast of West and South-West Europe. 

Primary moult timing and strategy

By October the majority of adult Lesser Black-backed Gulls moulting the outer primaries. There is a clear difference between presumed birds lingering around and birds which will probably migrate further south quickly. The difference can be found in the primary moult strategy. 

Some adult have a large moulting gap in the central primaries (for example P8-P10 old and only P4 fully grown and P5 still slightly shorter than P4) by the first days of September. It is hard to believe these birds will travel large distances non-stop as such moult gaps are highly inefficient for this migration pattern. Probably these birds, which can be found moulting in a period roughly from late August to late November stay in the area throughout their primary moult. They migrate further south either forced by cold winter or when the primaries moult arrives at the far outer primaries.

The other group is represented by birds with a very small moult gap in the primaries or with a very low primary moult score, indicating that primary moult will probably suspended until arrival at the wintering grounds. 

Tables & Graph

starting date:   24 Sep   01 Oct   08 Oct   15 Oct   22 Oct   29 Oct   05 Nov
week no: 39 40 41 42 43 44 45
REM 0 0 2 97 111 113 122 2
REM 1 19 1 183 165 56 3 0
REM 2 85 2 217 175 55 9 0
REM 3 123 2 106 38 7 1 0
REM 4 44 0 39 11 2 0 0
REM 5 24 0 28 7 1 0 0
REM 6 9 0 29 5 0 0 0
REM 7 5 0 8 3 0 0 0
REM 8 1 0 9 1 0 0 0
REM 9 0 0 3 1 0 0 0
REM 10 0 0 3 1 0 0 0
0 0 0 0 0 0 0
n = 310 7 722 518 234 135 2
average P = 3.1 1.6 2.2 1.5 0.9 0.2 0.0
95% CID = 0.14 1.18 0.13 0.12 0.13 0.10 0.00
inverse = 1.97 2.45 1.96 1.96 1.97 1.98 12.71
Variantie (s2) = 1.57 1.62 3.30 1.81 0.97 0.33 0.00
SD (s) = 1.25 1.27 1.82 1.35 0.98 0.57 0.00
SE (sx) = 0.07 0.48 0.07 0.06 0.06 0.05 0.00
               
% class:              
0 0 29 13 21 48 90 100
1 6 14 25 32 24 2 0
2 27 29 30 34 24 7 0
3 40 29 15 7 3 1 0
4 14 0 5 2 1 0 0
5 8 0 4 1 0 0 0
6 3 0 4 1 0 0 0
7 2 0 1 1 0 0 0
8 0 0 1 0 0 0 0
9 0 0 0 0 0 0 0
10 0 0 0 0 0 0 0
               
  % control: 100 100 100 100 100 100 100

  Table above, data for the Netherlands (2000-2010), results for October (aggregated week): number of remaining old primaries in adults.
REM0 = 0 old primaries left (P10 dropped).

Above: Scatter Plot for PMS in adult Lesser Black-backed Gulls. Number of remaining old primaries throughout the year (arrested moult excluded). Sample size is 6.171 birds.

Above: Scatter Plot for PMS in adult Lesser Black-backed Gulls. Number of remaining old primaries for October (arrested moult excluded). Sample size for this graph is 40 ringed birds + 1888 unringed birds.

starting date:

24 Sep

08 Oct

15 Oct

22 Oct

29 Oct

05 Nov

week no:

39

41

42

43

44

45

NEW 0

0

0

0

0

0

0

NEW P1

0

0

0

0

0

0

NEW P2

0

0

0

0

0

0

NEW P3

0

0

0

0

1

1

NEW P4

4

0

0

2

1

1

NEW P5

10

1

2

17

5

6

NEW P6

4

12

94

62

10

13

NEW P7

0

5

127

162

96

115

NEW P8

0

0

24

136

164

145

NEW P9

0

0

0

0

71

18

NEW P10

 

 

 

 

2

13

 

 

 

 

 

 

 

n =

18

18

247

379

350

312

average NEW P =

5.0

6.2

6.7

7.1

7.8

7.6

95% CID =

0.34

0.27

0.08

0.09

0.10

0.11

inverse  =

2.11

2.11

1.97

1.97

1.97

1.97

Variantie (s2) =

0.47

0.30

0.42

0.74

0.83

0.91

SD (s) =

0.69

0.55

0.65

0.86

0.91

0.95

SE (sx) =

0.16

0.13

0.04

0.04

0.05

0.05

 

 

 

 

 

 

 

% class

 

 

 

 

 

 

0

0

0

0

0

0

0

1

0

0

0

0

0

0

2

0

0

0

0

0

0

3

0

0

0

0

0

0

4

22

0

0

1

0

0

5

56

6

1

4

1

2

6

22

67

38

16

3

4

7

0

28

51

43

27

37

8

0

0

10

36

47

46

9

0

0

0

0

20

6

10

0

0

0

0

1

4

 

 

 

 

 

 

 

% control:

100

100

100

100

100

100

 

  Table above, data for the Netherlands (2000-2010), results for October (aggregated week): number of new, fully grown primaries in adults.
NEW0 = no new primaries visible (arrested moult excluded in this analysis). So either P1 still old, or P1 dropped, but the very tip still not visible.
NEW1 = P1 longest new primary. Range between "at least the tip of P1 is visible" until "tip of P2 still shorter than P1".
NEW2 = P2 longest new primary. NEW3 = etc.

Above: Scatter Plot for PMS in adult Lesser Black-backed Gulls throughout the year. Longest new primary is classified "fully grown". Sample size is 2.969 birds. Birds scored on NEW primaries (arrested moult excluded).

Above: Scatter Plot for PMS in adult Lesser Black-backed Gulls. Scoring on longest new primary, which is classified "fully grown" for October (arrested moult excluded). Sample size for this graph is 58 ringed birds + 1266 unringed birds.

The earliest date for birds with missing primaries is 30 April 2006, which is not the earliest day in our complete sample. We scored 98 Lesser Black-backed Gulls prior to that date, some already in February. As can be expected, none of these was in primary moult, and this group was excluded in our further analysis as it will only result in a misfit for the onset of moult, using linear regression statistics.

The last scoring days were in November, with a larger sample (133 adults) on 04 November 2010. From that date we have the last birds with retaining old primaries (13 birds with 1, 2 or even 3 old primaries; 120 birds with no old primaries left). From other days in November, only 10 birds were scored (all ringed adults). They all had no old primaries left.

The first tine a new primary (P1) was visible, was on 08 May 2006. Prior to this date, birds were scored missing inner primaries, but the very tip was still not visible (and therefore not scored). As soon as the tip of P1 was visible, this was automatically scored "fully grown" For the other primaries to be scored fully grown, they had to exceed the previous primary in length (so P2 fully grown, only from when it is longer than P1).

Peter Stewart presents onset of primary moult by scores for caught adult Lesser Black-backed Gulls in the Severn Estuary, U.K., repeated in the table below. To read his results, see this PDF file.  

Table: the number of old remaining primaries present. Compare PMS of adult graellsii at half monthly intervals. Data from the Severn Estuary Gull Group by Peter Stewart: The primary moult of the Lesser Black-backed Gull (special publication No 1). Scores in the Netherlands can be influenced by unringed nominate fuscus, in autumn in particular.

 

Netherlands

Severn Estuary U.K.

 

PMS range

N:

Average PMS

PMS range

N:

Average PMS

1-15 apr

10

49

10

8-10

216

9,9

16-30 apr

10

47

10

10

197

10,0

1-15 may

8-10

849

9,8

8-10

167

9,9

16-31 may

7-10

380

9,7

7-10

76

9,5

1-15 jun

8-10

104

9,6

7-10

33

8,6

16-30 jun
7-10
253
9,7
6-10
65
8,2
1-15 jul
6-10
605
9,2
5-10
176
7,4
16-31 jul
-
-
-
4-8
44
6,4
1-15 aug
3-10
143
6,6
3-10
165
5,2
16-31 aug
2-10
1284
5,4
3-8
127
4,6
1-15 sep
1-10
300
3,6
0-7
234
3,3
16-30 sep
0-7
645
2,9
0-7
173
2,1
1-15 oct
0-9
733
2,2
0-4
328
0,9
16-31 oct
0-10
750
1,3
0-3
244
0,3
1-15 nov
0-3
138
0,2
0-3
604
0,1
16-30 nov
0
5
0
0-1
341
0,0

LINEAR REGRESSION

Onset of primary moult can best be illustrated by a sinusoid for the first few weeks and last weeks, but there appears to be linear dependency over much of the central moult period.

LINEAR REGRESSION FOR OLD REMAINING PRIMARIES

Complete sample: Linear regression (n = 6171, R2= 0.820, sign: P<0,000)
PMS = -0,381* WEEK + 17,967

Ringed adults: Linear regression (n = 2069, R2= 0.800, sign: P<0,000)
PMS = -0,325* WEEK + 16,270

Unringed adult: Linear regression (n = 4102, R2= 0.779, sign: P<0,000)
PMS = -0,513* WEEK + 23,031

LINEAR REGRESSION FOR NEW FULLY GROWN PRIMARIES

Complete sample: Linear regression (n = 2969, R2= 0.930, sign: P<0,000)
PMS = 0,302* WEEK - 5,933

Ringed adults: Linear regression (n = 1209, R2= 0.888, sign: P<0,000)
PMS = 0,252* WEEK - 4,747

Unringed adult: Linear regression (n = 1760, R2= 0.833, sign: P<0,000)
PMS = 0,442* WEEK - 11,906

Remaining old primaries for adult LBBG year 2001 (the Netherlands)
. 0 1 2 3 4 5 6 7 8 9 10 n: m:
Oct 8-12 101 136 173 83 30 22 25 7 8 3 3 591 2,17
(aggregated Maasvlakte & Westkapelle) n: total; m: average

 

Adult LBBG primary scores, Naaktstrand - Maasvlakte, the Netherlands, October 19 2002
. p8 old p9 old p10 old no old total old
p5 f.g. 2 - - - 2
p6 f.g. 3 68 20 - 91
p7 f.g. - 19 60 39 118
p8 f.g. - - 2 22 24
total f.g. 5 87 82 61 235

Notes:
1. f.g. (fully grown): including primaries which are not completely fully grown, but all primaries of which the tip exceeds the previous primary in the folded wing.
2. old primaries: total (n) = 235, average (m) = 8,8 standard deviation (SD) = 0,86.
3. new primaries: total (n) = 235, average (m) = 6,7 standard deviation (SD) = 0,66.
4.Adult LBBG scored in a group of 1.500 birds (of which 90% LBBG). A remarkable homogeneous group regarding grey-tone of upper-parts, after three days of western winds.
5. a few ringed birds in this group: 
orange CA9 (P10 old, P7 f.g.) ringed as pullus in the Netherlands on July 01 1995, now 8cy.
green EA34 (P10 old, P7 f.g.) ringed as pullus in the Netherlands on July 02 1993, now 10cy.
green EV62 (P10 old, P7 f.g.) ringed as pullus in the Netherlands on June 21 1997, now 6cy.

 

Adult LBBG primary scores, Dannes (region Calais), NW France, October 19 2002
. p6 old p7 old p8 old p9 old p10 old no old total old
p4 f.g. - 1 - - - - 1
p5 f.g. 1 1 5 1 - - 8
p6 f.g. - - 2 30 11 - 43
p7 f.g. - - - 23 37 30 90
p8 f.g. - - - 1 8 83 92
total f.g. 1 2 7 55 56 113 234

Notes:
1. f.g. (fully grown): including primaries which are not completely fully grown, but all primaries of which the tip exceeds the previous primary in the folded wing.
2. old primaries: total (n) = 234, average (m) = 9,1 standard deviation (SD) = 0,98.
3. new primaries: total (n) = 234, average (m) = 7,1 standard deviation (SD) = 0,86.
4.Adult LBBG scored in a group of 2.000 birds (of which 30% LBBG). A heterogeneous group regarding grey-tone of upper-parts.
5. a few ringed birds in this group: 
orange EV89 (P8 f.g.) ringed as pullus in the Netherlands on July 02 1997, now 6cy.
green E774 (P9 = P8 f.g.) ringed as pullus in the Netherlands on July 04 1995, now 8cy.
white R23 (P9 old, P6 f.g.) ringed as pullus in the Netherlands on July 06 1991, now 12cy.
blue ASAY (P8 f.g.) ringed in Belgium.

.
Britain: 'graellsii'
.
ad LBBG in October, ringed in the U.K. (153884 bytes)graellsii ad APF October 21 2004, Westkapelle, the Netherlands. Picture Pim Wolf.
ad LBBG in October, ringed in the U.K. (57532 bytes)graellsii ad ATN October 22 2004, Westkapelle, the Netherlands. Picture Pim Wolf.
ad LBBG in October, ringed in the U.K. (61530 bytes)graellsii 6cy LSY October 22 2004, Westkapelle, the Netherlands. Picture Pim Wolf.
ad LBBG in October, ringed in the U.K. (115719 bytes)graellsii ad YLS October 19 2004, Westkapelle, the Netherlands. Picture Pim Wolf.
ad LBBG in October, ringed in the U.K. (122109 bytes)graellsii adGG-63238Westkapelle, the Netherlands, October 04 2004. Photo by Pim Wolf.
.
Belgium: 'Dutch intergrade'
.
ad LBBG in October, ringed in the Netherlands. (60900 bytes)Dutch intergrade 15cy & 17cy DKAN October 2009 & September 2011, Westkapelle, the Netherlands.
ad LBBG in October, ringed in Belgium. (74645 bytes)Dutch intergrade BLB L - 7400? October 05 2002, Le Portel / Boulogne-sur-Mer, NW France.
ad LBBG in October, ringed in Belgium. (85453 bytes)Dutch intergradeBLB L - 80954 October 22 2001, Westkapelle, the Netherlands.
ad LBBG in October, ringed in Belgium. (75750 bytes)Dutch intergrade 5cy BLB L - 85168 October 30 2003, Westkapelle, the Netherlands.
.
the Netherlands: ' Dutch intergrade'

.
juvenile LBBG, ringed in the Netherlands. LBBG RD October 11 2010, Madrid, Spain. Photo by Delfín González.
juvenile LBBG, ringed in the Netherlands. LBBG WE October 11 2010, Madrid, Spain. Photo by Delfín González.
juvenile LBBG, ringed in the Netherlands. LBBG HF October 11 2010, Madrid, Spain. Photo by Delfín González.
juvenile LBBG in October, ringed in Norway. (131273 bytes)LBBG P8 October 27 2006, Westkapelle, the Netherlands.
juvenile LBBG, ringed in the Netherlands. LBBG 6L October 08 2014 & May 2006, the Netherlands & France. Picture: Jean-Michel Sauvage.
2275.jpg (40492 bytes) LBBG 6cy KLAX October 21 2012, Spain. Also as 4cy in Holland.
LBBG YAHJ 2009-2011, France & the Netherlands. Pictures: Ruud Altenburg & Jean-Michel Sauvage.
adult LBBG Dutch intergrade in September, ringed in the Netherlands. (80494 bytes)LBBG YAKZ 2010-2011, IJmuiden-Westkapelle, the Netherlands.
ad LBBG in October, ringed in the Netherlands. (77525 bytes)15cy E864 October 06 2009, Westkapelle, the Netherlands.
ad LBBG in October, ringed in the Netherlands. (77525 bytes)5cy ECJ9 October 14 2001, Westkapelle, the Netherlands.
ad LBBG in October, ringed in the Netherlands. (85975 bytes)9cy EN22 October 19 2004, Westkapelle, the Netherlands.
ad LBBG in October, ringed in the Netherlands. (86535 bytes)11cy JP2 October 11 2003, Westkapelle, the Netherlands.
ad LBBG in October, ringed in the Netherlands. (60900 bytes)16cy JR6 October 19 2008, Madrid, Spain. Photo by Delfín González.
ad LBBG in October, ringed in the Netherlands. (60900 bytes)11cy KN8 October 03 2003, Scheveningen, the Netherlands.
ad LBBG in October, ringed in the Netherlands. (65112 bytes)10cy E737 October 04 2004, Westkapelle, the Netherlands. Picture Pim Wolf.
ad LBBG in October, ringed in the Netherlands. (67338 bytes)8cy EL74 October 11 2003, Westkapelle, the Netherlands.
ad LBBG in October, ringed in the Netherlands. (73481 bytes)6cy ECK1October 03 2002, Boulogne-sur-Mer, NW France.
ad LBBG in October, ringed in the Netherlands. 6cy EH10Westkapelle, the Netherlands, October 21 1999. Photo by Pim Wolf. Born summer 1994 in the Netherlands.
ad LBBG in October, ringed in the Netherlands. 7+cy ER73Westkapelle, the Netherlands, October 15 & 27 1999. Photo by Pim Wolf.
ad LBBG in October, ringed in the Netherlands. (125356 bytes)5cy E214 October 04-07 2004, Westkapelle, the Netherlands. Picture Pim Wolf.
ad LBBG in October, ringed in the Netherlands. (142636 bytes)5cy E219 October 01 2004, Westkapelle, the Netherlands. Picture Pim Wolf. Also seen as 2cy at Westkapelle.
ad LBBG in October, ringed in the Netherlands. (142636 bytes)5cy 7F October 09 2010, Heemskerk, the Netherlands.
ad LBBG in October, ringed in the Netherlands. (86318 bytes)5.362.926 October 13 2001, Westkapelle, the Netherlands.
juvenile LBBG in October, ringed in Norway. (131273 bytes)5.366.905 October 09 2005, Westkapelle, the Netherlands.

.N Germany: transition
graellsii - intermedius
.
2cy intermedius in October, ringed in Norway. (97235 bytes)H.501 September - October 2010, Leiden, the Netherlands.
2cy intermedius in October, ringed in Norway. (97235 bytes)H.595 October 06 2009, Westkapelle, the Netherlands.
ad LBBG in winter, ringed in the Netherlands. (65647 bytes)LBBG (graellsii) 7cy DEH 4290815 October 03 2012, Boulogne-sur-Mer, NW France. Picture: Jean-Michel Sauvage.
.Denmark: intermedius
.
2cy intermedius in July, ringed in Norway. (114585 bytes)6cy LBBG intermediusV3SJ July 2010, August 2014 & October 2014, Utrecht, the Netherlands. Also as 2CY.
adult LBBG intermedius in September, ringed in Norway. (95609 bytes)LBBG VC5M October 24 2012, Playa de Pantin - Valdovino, Spain (43.38N-008.07W). Picture: Antonio Gutierrez.
adult intermedius in October, ringed in Denmark. (83054 bytes)intermedius 9cy 467213 October 14 2001, Westkapelle, the Netherlands. Ringed in Denmark: Veno, Struer, Jylland (56.33N 08.37E).
adult intermedius in October, ringed in Denmark. intermedius 28cy 4041636Weskapelle, the Netherlands, October 27 1999. Ringed at Anholt, Denmark in 1972. Photo by Pim Wolf.
adult intermedius in October, ringed in Denmark. intermedius 26cy 4041916Westkapelle, the Netherlands, October 26 1999. Photo by Pim Wolf.
adult intermedius in October, ringed in Denmark. intermedius 13cy 4124822Westkapelle, the Netherlands, October 25-27 1999. Photo by Pim Wolf.
adult intermedius in October, ringed in Denmark. (71780 bytes)intermedius4130670Westkapelle, the Netherlands, October 04 2004. Photo by Pim Wolf.
adult intermedius in October, ringed in Denmark. (72633 bytes)intermedius4253409IJmuiden & Westkapelle, the Netherlands, October 2003 & October 2004.
adult intermedius in October, ringed in Denmark. (112876 bytes)intermediusxxx Westkapelle, the Netherlands, October 19 2004. Photo by Pim Wolf.

.Norway: intermedius
.
adult intermedius lbbg in October, ringed in Norway. (94818 bytes)intermedius 5cy JJ5A October 09 2009, IJmuiden, the Netherlands.
adult intermedius lbbg in October, ringed in Norway. (94818 bytes)intermedius 5cy+ JM79 October 19 2002, Westkapelle, the Netherlands.
adult intermedius lbbg in October, ringed in Norway. (65103 bytes)intermedius JS26 October 10 2004, Westkapelle, the Netherlands. Picture Pim Wolf.
juvenile LBBG in October, ringed in Norway. (131273 bytes)intermedius 5+cy J5YL October 14 2012, Westkapelle, the Netherlands.
juvenile LBBG, ringed in the Netherlands. LBBG adult JE8J October & December 2012, Madrid, Spain. Photo by Delfín González.
adult intermedius lbbg in October, ringed in Norway. (90595 bytes)intermedius 41699 October 22 2001, Westkapelle, the Netherlands. Ringed Stavanger, Norway.
juvenile LBBG in October, ringed in Norway. (131273 bytes)intermedius 25cy 492441 October & November, Westkapelle, the Netherlands. Picture Pim Wolf.
adult intermedius lbbg in October, ringed in Norway. (81887 bytes)intermedius 18cy 4100410 October 11 2003, Westkapelle, the Netherlands.
adult intermedius lbbg in October, ringed in Norway. (55754 bytes)intermedius 4105805IJmuiden, the Netherlands, October 02 2003.
adult intermedius lbbg in October, ringed in Norway.intermedius 14cy 4108778Westkapelle, the Netherlands, October 27 1999. Photo by Pim Wolf.
adult intermedius lbbg in October, ringed in Norway. (137939 bytes)intermedius4119689October 19 2004, Westkapelle, the Netherlands. Picture Pim Wolf.
adult intermedius lbbg in October, ringed in Norway. (75971 bytes)intermedius4175071 October 19 2004, Westkapelle, the Netherlands.
adult intermedius lbbg in October, ringed in Norway. (74314 bytes)intermedius4178002October 21 2004, Westkapelle, the Netherlands. Picture Pim Wolf.
adult intermedius lbbg in October, ringed in Norway. (99830 bytes)intermedius4199556October 21 2004, Westkapelle, the Netherlands. Picture Pim Wolf.
adult intermedius lbbg in October, ringed in Norway. (62093 bytes)intermediusNOO NA 08134 October 19 2004, Westkapelle, the Netherlands.
adult intermedius lbbg in October, ringed in Norway. (51593 bytes)intermediusNOO NA 0846? October 10 2003, Scheveningen, the Netherlands. Ringed Oslo, Norway.
adult intermedius lbbg in October, ringed in Norway. (77684 bytes)intermediusNOO NA 09175 October 18 2002, Westkapelle, the Netherlands. Ringed Oslo, Norway.

.Sweden: intermedius
.
adult intermedius lbbg in October, ringed in Sweden. (80211 bytes)intermedius 15CY SVS 8072924 October 19 2004, Westkapelle, the Netherlands.
adult intermedius lbbg in October, ringed in Sweden. (102169 bytes)intermedius LBBG SVS 8080900 October 19 2004, Westkapelle, the Netherlands.
adult intermedius lbbg in October, ringed in Sweden. (102169 bytes)intermedius LBBG SVS 8088508 October 22 2009, Westkapelle, the Netherlands.
adult intermedius lbbg in October, ringed in Sweden. (102169 bytes)intermedius LBBG SVS 8105583 October 06 2009, Westkapelle, the Netherlands.
.
Ringed adultsintermedius (no ring-data received yet)
.
5488int04546.jpg (63731 bytes)LBBG intermedius ?04546, October 04 2002, Le Portel / Boulogne-sur-Mer, NW France.
LBBG intermedius adult 408972, Breskens, the Netherlands, October 20 1999. Photo by Pim Wolf. Most probably with Stavanger (Norway) ring.
LBBG intermedius adult 492155, Breskens, the Netherlands, October 20 1999. Photo by Pim Wolf. Most probably with Stavanger (Norway) ring.
LBBG intermedius adult ..746.., Breskens, the Netherlands, October 20 1999. Photo by Pim Wolf.
5491int4196602.jpg (58985 bytes)LBBG intermedius 4196602, October 04 2002, Le Portel / Boulogne-sur-Mer, NW France.
1cy argenteus in November, ringed in Belgium. (25637 bytes)LBBG adult N537 October 05 2005, Westkapelle, the Netherlands.
.
Unringed adults in October
.
7435.jpg (59166 bytes)Link to 13 unringed adult LBBG, October 22 2001, Westkapelle, the Netherlands.